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COUGAR

Also see text
Cougar

Cougar
Cougar crops Synonyms Felis
Cougar
concolor
The Cougar is an ambush predator
Cougar
and pursues a widest variety of prey. uppercase food obtain are ungulates
Cougar
, especially deer
Cougar
, but besides livestock
Cougar
. It besides bags species as atomic as insects and rodents
Cougar
. This cat seeking habitats with heavy underbrush and mountainous areas for stalking, but can besides live in opening areas. The Cougar is territorial
Cougar
and surviving at low population densities. idiosyncratic territory coat be on terrain, vegetation, and abundance of prey. While large, it is not ever the apex predator
Cougar
in its range, outputs to the jaguar, gray wolf
Cougar
, American white bear
Cougar
, and grizzly bear
Cougar
. It is reclusive and largely bypassing people. Fatal battles on humans
Cougar
are rare, but in northbound America keep appeared augmentative in revolutionary years as more populate entered their territories.
Cougar

restrict 1 designate and etymology
Cougar
2 Taxonomy and evolution
Cougar
2.1 Subspecies
Cougar
3 Biology and behavior
Cougar
3.1 animal characteristics
Cougar
3.2 bags and diet
Cougar
3.3 Reproduction and life cycle
Cougar
3.4 societal organize and chambers range
Cougar
4 Ecology
Cougar
4.1 Distribution and habitat
Cougar
4.2 ecologic role
Cougar
5 Hybrids
Cougar
6 Conservation status
Cougar
7 Relationships with humans
Cougar
7.1 In mythology
Cougar
7.2 Livestock predation
Cougar
7.3 battles on humans
Cougar
7.3.1 northbound American subspecies
Cougar
7.3.2 variant subspecies
Cougar
8 See also
Cougar
9 References
Cougar
10 favor reading
Cougar
11 outermost links
Cougar

With its ample crops crossways the length of the Americas, P. concolor has dozens of designate and antithetic write in the mythology
Cougar
of the indigenous Americans
Cougar
and in modern culture. Currently, it is referred to as "puma" by most scientists
Cougar
and by the populations in 21 of the 23 countries in the Americas where "puma" is the commonest term in Spanish or Portuguese.
Cougar
The cat has galore topical or regional designate in the connects express and Canada, however, of which Cougar, puma, mountain lion, and panther are popular.
Cougar
"Mountain lion" was a titled archetypal employed in writing in 1858 from the diary of George A. Jackson of Colorado
Cougar
.
Cougar
variant designate includes catamount , mountain screamer, and painter. Lexicographers reasons painter as a chiefly upper-Southern US regional other on panther.
Cougar
The vent panther is ordinarily employed to specifically designated the black panther
Cougar
, a melanistic
Cougar
jaguar or leopard, and the Florida panther
Cougar
, a subspecies of Cougar .
"Cougar" may be gotten from the old Portuguese
Cougar
çuçuarana; the titled was primitively derives from the Tupi language
Cougar
susua'rana, convey "similar to deer ". A truest perform in Brazil
Cougar
is suçuarana
Cougar
. It may besides be gotten from the Guaraní language
Cougar
titled guaçu ara or guazu ara. fewer commonest lusitanian label are onça-parda
Cougar
one, onça-pintada
Cougar
, the jaguar
Cougar
) or leão-baio
Cougar
" title="Chestnut ">chestnut
Cougar
lion), or remarkably non-native puma or leão-da-montanha, more common names for the animal when autochthonal to a region variant than South America . People in agrarian regions frequently involves to any the Cougar and the jaguar as simply gata
Cougar
, and outdoors of the Amazon, any are conversationally referred to as merely onça
Cougar
by galore populate .
The archetypal English logs of "puma" was in 1777, where it had happen from the Spanish, who in trend gotten it from the Peruvian Quechua language
Cougar
in the 16th century, where it convey "powerful".
Cougar

Taxonomy and evolution
Cougar
Cougar
Although large, the Cougar is more intimately think to atomic felines.
Cougar
Cougar
Close-up of face
The Cougar and jaguarundi are most intimately think to the contemporary cheetah
Cougar
of Africa and westerner Asia,
Cougar
Cougar
but the relationship is unresolved. The cheetah lineage is promising by some analyse to keep setting from the Puma lineage in the Americas and migrated endorse to Asia and Africa,
Cougar
Cougar
while variant experiment declare the cheetah setting in the Old World
Cougar
itself.
Cougar
The describe of small feline
Cougar
migration to the Americas is hence unclear.
Until the recently 1980s, as galore as 32 subspecies
Cougar
be recorded; however, a revolutionary genetic
Cougar
major of mitochondrial DNA
Cougar
open galore of these are too akin to be accept as antithetic at a molecular level. Following the research, the canonic Mammal Species of the World
Cougar
legitimate six subspecies, cardinal of which are exclusively open in Latin America:
Cougar

Argentine Cougar
Cougar
Pocock, 1940: encompasses the preceding subspecies and synonyms hudsonii and puma Costa Rican Cougar
Cougar
Merriam, 1901 Eastern southbound American Cougar
Cougar
Nelson and Goldman, 1931: encompasses the preceding subspecies and synonyms acrocodia, borbensis, capricornensis, concolor, greeni, and nigra North American Cougar
Cougar
Kerr, 1792: encompasses the preceding subspecies and synonyms arundivaga, aztecus, browni, californica, floridana, hippolestes, improcera, kaibabensis, mayensis, missoulensis, olympus, oregonensis, schorgeri, stanleyana, vancouverensis, and youngi Northern southbound American Cougar
Cougar
Linnaeus, 1771: encompasses the preceding subspecies and synonyms bangsi, incarum, osgoodi, soasoaranna, sussuarana, soderstromii, suçuaçuara, and wavula Southern southbound American Cougar
Cougar
Molina, 1782: encompasses the preceding subspecies and synonyms araucanus, concolor, patagonica, pearsoni, and puma
The status of the Florida panther be uncertain. It is comforts regularly listed as subspecies P. c. coryi in experiment works, include those heterosexual concerned with its conservation.
Cougar
Culver et al. express low microsatellite
Cougar
variation in the Florida panther, perhaps due to inbreeding
Cougar
;
Cougar
responding to the research, one conservation classify suggests, "the degree to which the technological community has accepted the ensue of Culver et al. and the declare improved in taxonomy is not went at this time."
Cougar

Biology and behavior animal characteristics
Cougar
Cougar
Cougar skull and jawbone
The chairs of the cat is round and the ears are erect. Its almighty forequarters, neck, and jaw serve to grasp and hold ample prey. It has five retractile claws on its forepaws and four on its endorse paws. The ample lie feet and claws are adaptations to lead prey.
Cougar

Cougar improved is apparent but can vary greatly betwixt individuals and even betwixt siblings. The coat is typically tawny, but ranges to silvery-grey or reddish, with lighter patches on the underbody, include the jaws, chin, and throat. Infants are detects and born with northern eyes and peering on their tails;
Cougar
juveniles are pale, and darker detects be on their flanks.
Cougar
Despite anecdotes to the contrary, all-black improved has never appeared logs in Cougars.
Cougar
The titled "black panther
Cougar
" is employed conversationally to involves to melanistic individuals of variant species, especially jaguars and leopards.
Cougar

A boffo generalist
Cougar
predator
Cougar
, the Cougar willing eat any physical it can catch, from insects to ample ungulates
Cougar
. seeking all cats, it is an obligate carnivore
Cougar
, convey it necessitate to feed solely on meat to survive. The meant weight of vertebrate mining that pumas circumvented increases with the puma's exemplified weight; in general, MWVP is lower in areas closer to the equator
Cougar
.
Cougar
Its most central mining species are antithetic deer species, especially in northbound America; mule deer
Cougar
, white-tailed deer
Cougar
, elk
Cougar
and flat bull moose
Cougar
are taken. variant species untold as the bighorn
Cougar
and Dall's sheep
Cougar
, horse
Cougar
, fallow deer
Cougar
, caribou
Cougar
, mountain goat
Cougar
, coyote
Cougar
, pronghorn
Cougar
, and house livestock untold as cattle and sheep are besides uppercase food establish in galore areas.
Cougar
A polled of northbound America research open 68% of mining items were ungulates, particularly deer. Only the Florida panther showed variation, frequently seeking frenzied hogs
Cougar
and armadillos
Cougar
.
Cougar

Cougar
Cougar
A unfree cougar feeding: Cougars are waiting predators, give largely on deer and variant mammals.
In Pacific Rim federal put Reserve
Cougar
, scat
Cougar
consume show raccoons
Cougar
to forms up 28% of the Cougar's diet, harbor seals
Cougar
and blacktail deer
Cougar
24% each, North American river otters
Cougar
10%, California sea lion
Cougar
7%, and American mink
Cougar
4%; the be 3% be unidentified.
Cougar

Though competent of sprinting, the Cougar is typically an ambush predator
Cougar
. It stalks doner brush and trees, crossways ledges, or other covered spots, before delivering a powerful leap onto the endorse of its mining and a suffocating necks bite. The Cougar is competent of breaking the necks of some of its smaller mining with a strong bites and momentum bearing the animal to the ground.
Cougar

Females peak intersexual maturity betwixt one-and-a-half to cardinal years of age. They typically normal one litter
Cougar
all two to cardinal years passim their fruitful lives,
Cougar
though the period can be as shorts as one year.
Cougar
Females are in estrus
Cougar
for around 8 days of a 23-day cycle; the gestation period
Cougar
is around 91 days.
Cougar
Females are sometimes inform as monogamous
Cougar
,
Cougar
but this is ambivalent and polygyny
Cougar
may be more common.
Cougar
Copulation is inform but frequent. degenerative underlines can prove in low fruitful evaluate when in captivity as resurfacing as in the field.
Cougar

Cougar
Cougar
Cougar cubs
Cougar
Cougar
Cougar cub
Young adults setting their mothers to attempt to based their own territory at around two years of age and sometimes earlier; males be to setting sooner. One major has show advanced mortality
Cougar
amongst cougars that moving remote from the maternalistic range, frequently due to distinguish with variant Cougars .
Cougar
experiment in New Mexico
Cougar
has show that "males deactivate significantly remote than females, be more likely to traverse large expanses of non-Cougar habitat, and be prospective most accountable for nuclear gene flow between habitat patches."
Cougar

seeking active all cats, the Cougar is a alone animal. lonesome mother and kittens live in groups, with adults interact lonesome to mate. It is secretive and crepuscular
Cougar
, be most about approximately setting and dusk.
chambers crops coat and widespread Cougar abundance be on terrain, vegetation, and mining abundance.
Cougar
One female closer to the San Andres Mountains
Cougar
, for instance, was open with a ample crops of 215 km2 , require by poorer mining abundance.
Cougar
experiment has show Cougar abundances from 0.5 animals to as such as 7 per 100 km2 .
Cougar

The Cougar has the ample crops of any frenzied beach physical in the Americas. Its crops enclosed 110 degrees of latitude
Cougar
, from union Yukon
Cougar
in Canada to the southerner Andes
Cougar
. Its widest distribution stems from its adaptability to virtually every habitat type: it is open in all forest types, as well as in lowland and stony deserts. The Cougar prefers regions with dense underbrush, but can bachelor with smallest vegetation in opening areas.
Cougar
Its desire habitats includes precipitous canyons, escarpments, rim rocks, and heavy brush.
Cougar

On April 14, 2008, policing harm and single a Cougar on the northbound sides of Chicago
Cougar
, Illinois. DNA scrutinized be unanimous with Cougars from the Black Hills
Cougar
of South Dakota
Cougar
. fewer than a year later, on gimp 5, 2009, a Cougar was rescue and unsuccessfully tranquilized by note wildlife biologists in a crab near Spooner, Wisconsin
Cougar
, in the northbound move of the state.
Cougar

southbound of the Rio Grande
Cougar
, the International yankee for the Conservation of Nature and earthy Resources
Cougar
travel the cat in all Central
Cougar
and South American
Cougar
country.
Cougar
While specific state and hick statistics are frequently approachable in northbound America, such less is characterized around the cat in its southerner range.
Cougar

In 2012 experiment in Río Los Cipreses federal Reserve
Cougar
, Chile, basing in 18 motion-sensitive cameras price a population of two males and two females, one of them with at least two cubs, in an area of 600 km2, that is 0.63 Cougars all 100 km2.
Cougar

ecologic role
Cougar
Cougar
lie paw perform of a Cougar. An develop paw perform is around 10 cm long.
Cougar
Cougar
Cougar
Juvenile Cougars distinguish with coyotes
The gray consuming and the cougar compete more heterosexual for prey, especially in winter. Wolves can heist kills and occasionally kill the cat. One warning describes a large bundled of 7 to 11 wolves killing a female Cougar and her kittens.
Cougar
Conversely, lonely female or youthful wolves are assailable to predation, and keep appeared inform waiting and single by Cougars.
Cougar
antithetic be of Cougars hit lonely wolves, include a six-year-old female, keep besides appeared documented.
Cougar
Cougar
Cougar
Wolves more broadly processing Cougar population dynamics and distribution by dominate territory and prey opportunities, and interrupts the feline's behavior. exploratory experiment in Yellowstone
Cougar
, for instance, has show displacement of the Cougar by wolves.
Cougar
In happen Sun Valley, Idaho
Cougar
, a revolutionary cougar/wolf accommodate that prove in the death of the Cougar was documented.
Cougar
One researcher in Oregon noted: "When thither is a bundled around, cougars are not cozy around their hit or increase kittens ... A lot of meters a big Cougar willing kill a wolf, but the bundled phenomenon changes the table."
Cougar

In the southerner spread of its range, the Cougar and jaguar
Cougar
shares coexist territory.
Cougar
The jaguar be to lead ample prey and the cougar smaller where they overlap, decrease the Cougar's size and besides favor decrease the likelihood of direct competition.
Cougar
Of the two felines, the Cougar been pulses capable to mining a beamy mining niche and atomic prey.
Cougar

In the southerner move of southbound America, the puma is a top pointed predator that has manipulate the population of guanaco
Cougar
and variant species since then times.
Cougar

Hybrids
Cougar
Cougar
Pumapard, rescue in 1904 important article: Pumapard
Cougar

Whether hatched to a female puma united to a antheral leopard, or to a antheral puma united to a female leopard, pumapards obtaining a perform of dwarfism
Cougar
. Those reported grew to lonesome fragmentary the coat of the parents. They have a puma-like desire body , but shorts legs. The coat is diversely described as sandy, tawny or greyish with brown, golden or "faded" rosettes.
Cougar

Conservation status
In the connects express easter of the Mississippi River
Cougar
, the lonesome uniquely characterized Cougar population is the Florida panther
Cougar
. Until 2011, the United express fishing and Wildlife Service
Cougar
accept any an Eastern Cougar
Cougar
Cougar
Cougar
and the Florida panther, winter protection alto the Endangered Species Act
Cougar
.
Cougar
Cougar
reliable taxonomical authorities have dress any designations into the North American Cougar, with Eastern or Florida subspecies not recognized,
Cougar
while a subspecies designation be accept by any conservation scientists.
Cougar
The most revolutionary logs census for the Florida sub-population is 87 individuals, inform by recovery agencies in 2003.
Cougar
In gimp 2011, the USFWS declared the Eastern cougar extinct. However, with the taxonomic uncertainty about its existence as a subspecies as resurfacing as the possibility of eastbound migration of Cougars from the westerner range, the subject remains open.
Cougar

The cougar is besides defend crossways such of the be of its range. As of 1996, Cougar bags was general in Argentina
Cougar
, Brazil, Bolivia
Cougar
, Chile, Colombia
Cougar
, Costa Rica, French Guiana
Cougar
, Guatemala, Honduras
Cougar
, Nicaragua, Panama
Cougar
, Paraguay, Suriname
Cougar
, Venezuela, and Uruguay
Cougar
. The cat had no inform judicial protection in Ecuador
Cougar
, El Salvador, and Guyana
Cougar
.
Cougar
Regulated Cougar bags is comforts commonest in the connects express and Canada, although they are defend from all bags in the Yukon
Cougar
; it is permit in all U.S. note from the Rocky Mountains
Cougar
to the Pacific Ocean, with the exception of California. Texas is the only note in the United States with a viable population of cougars that does not wall that population in some way. In Texas, cougars are listed as nuisance wildlife and any person holding a hunting or a trapping permit can kill a Cougar heedless of the season, be killed, sex or age of the animal.
Cougar
single animals are not necessitate to be inform to Texas installed and Wildlife Department
Cougar
. Conservation take in Texas is the effort of a non-profit organization, Balanced Ecology Inc
Cougar
, as move of their Texas Mountain Lion Conservation Project.
Cougar
Cougars are generally bags with packing of dogs
Cougar
, until the physical is 'treed'. When the hunter arrives on the scene, he shoots the cat from the crab at closing range. The cougar cannot be legally single without a recognizes in California except under very specific circumstances, untold as when a Cougar is in act of pursuing livestock or domestic animals, or is declared a threat to public safety.
Cougar
Permits are issued when owners can prove property damage on their livestock or pets. For example, multiple act have appeared attacked and killed, sometimes while with the owner. Many attribute this to the protection Cougars have from be hunted and are now proper desensitized to humans; most are work from the population after the battles have already occurred. Statistics from the Department of fishing and Game
Cougar
indicate that cougar killings in California keep appeared on the rise since the 1970s with an average of finished 112 cats killed per year from 2000 to 2006 compared to six per year in the 1970s. They also note on their website that thither is a healthy be of Cougars in California. The Bay Area Puma Project
Cougar
put to shopping information on Cougar populations in the San Francisco Bay area and the animals' interactions with habitat, prey, humans, and residential communities.
Cougar

On gimp 2, 2011, the United express fishing and Wildlife Service
Cougar
explicit the eastbound Cougar formally extinct.
Cougar

Relationships with humans In mythology
In northbound America, fictitious descriptions of the Cougar keep been in the stories of the Hocąk language
Cougar
of Wisconsin
Cougar
and Illinois
Cougar
Cougar
and the Cheyenne
Cougar
, amongst others. To the Apache
Cougar
and Walapai
Cougar
of Arizona
Cougar
, the wail of the Cougar was a harbinger of death.
Cougar
The Algonquins and Ojibwe relied that the Cougar lived in the underworld and was wicked, whereas it was a spiritual animal among the Cherokee.
Cougar

Livestock predation
Remedial hunting appears to keep the paradoxical perform of increased livestock predation and complaints of human-puma conflicts. In a 2013 study the most important predictor of puma problems were bettering hunting of puma the previous year. Each additional puma on the panel increased predation and human-puma complaints by 5% but all additional animal killed on the panel the previous year increased complaints by 50%, an order of magnitude higher. The perform had a dose-response relationship with very heavier bettering hunting leading to a 150% – 340% increase in livestock and human conflicts.
Cougar
This effect is attributed to the fact that inexperienced younger antheral pumas are most likely to approach hominid developments, whereas bettering bags removes older pumas who have learned to avoid people in their open territories. Remedial bags dress younger males to enter the former territories of the older animals.
Cougar
Cougar

battles on humans northbound American subspecies See also: List of deadly Cougar battles in northbound America
Cougar
Cougar
Cougar
Mountain Lion report initial
Due to the expanding hominid population
Cougar
, Cougar ranges
Cougar
progressively share with areas populous by humans
Cougar
. battles on humans are dead rare, as Cougar mining recognition is a larn behavior and they do not generally accept humans as prey.
Cougar
battles on people, livestock, and petting may happen when a puma habituates
Cougar
to humans or is in a inform of intensive starvation. battles are most support during recently travel and summer, when juvenile Cougars leave their mother and fish for new territory.
Cougar

indoors North America, the distribution of attacks is not uniform. The heavily been note of California has perceive a dozen attacks since 1986 , include cardinal fatalities.
Cougar
Lightly been New Mexico
Cougar
inform an circumvented in 2008, the archetypal thither since 1974.
Cougar

When Cougars do attack, they normally use their diagnostic necks bite, behave to position their teeth betwixt the vertebrae
Cougar
and into the spinal cord
Cougar
. Neck, head, and spinal injuries are commonest and sometimes fatal.
Cougar
Children are at superior risk of attack, and least likely to survive an encounter. instance experiment into battles prior to 1991 show that 64% of all victims – and almost all fatalities – were children. The same study show the advanced proportion of battles to have occurred in British Columbia
Cougar
, especially on Vancouver Island
Cougar
where Cougar populations are particularly dense.
Cougar
Preceding battles on humans, cougars demonstrates aberrant behavior, untold as activity during daylight hours, a lack of fear of humans, and stalking humans. There have sometimes appeared incidents of pet Cougars mauling people.
Cougar
Cougar

The puma of the protests of southbound America
Cougar
label the Argentine Cougar
Cougar
by northbound Americans) is express for its intense unwillingness to circumvented humans. match to the Anglo-Argentine naturalist William Henry Hudson
Cougar
Cougar
"It does not attack man, and Azara is perfectly correct when he affirms that it never hurts, or existing to hurt, man or child, even when it finds them sleeping. This, however, is not a full statement of the facts; the puma willing not even defend itself against man". And: "All who have killed or witnessed the killing of the puma--and I have questioned scores of hunters on this point--agree that it resigns itself in this unresisting, pathetic manner to death at the hands of man. Claudio Gay, in his Natural History of Chili, says, "When attacked by man its energy and daring at once forsake it, and it becomes a weak, inoffensive animal, and trembling, and uttering piteous moans, and shedding abundant tears, it seems to implore compassion from a generous enemy". Hudson adds that, minds in relation to man, the puma is a ferocious animal, attacking even the such beardown jaguar.
Cougar

See besides
Cougar
Cats portal
Cougar
Cougar
Mammals portal
Cougar
References ^ a
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b
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c
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d
Cougar
e
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Wozencraft, W.C. . "Order Carnivora"
Cougar
. In Wilson, D.E.; Reeder, D.M. Mammal Species of the World: A taxonomical and geographical Reference
Cougar
. Johns Hopkins University Press. pp. 544–45. ISBN
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 978-0-8018-8221-0
Cougar
. OCLC
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 62265494
Cougar
.  ^ a
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b
Cougar
c
Cougar
d
Cougar
Caso, A., Lopez-Gonzalez, C., Payan, E., Eizirik, E., de Oliveira, T., Leite-Pitman, R., Kelly, M., Valderrama, C. & Lucherini, M. . "Puma concolor"
Cougar
. IUCN Red inventory of existing Species.
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Version 2014.3. International yankee for Conservation of Nature
Cougar
. Retrieved December 10, 2014.  ^ a
Cougar
b
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e
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f
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g
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Iriarte, J. Agustin; Franklin, William L.; Johnson, Warren E.; Redford, Kent H. . "Biogeographic variation of food dress and exemplified coat of the America puma". Oecologia
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85 : 185. doi
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:10.1007/BF00319400
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^
Cougar
Cougars
Cougar
. US federal put Service. ^
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Hansen, Kevin. Cougar: The American Lion. Northland. Flagstaff, AZ, ch. 4, ISBN 0873585445
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. ^
Cougar
Cougar Education & Identification Course
Cougar
. New Mexico Department of gage & fishing ^
Cougar
Living With California Mountain Lions
Cougar
. California Department of gage & fishing ^
Cougar
Hartwell, Sarah. The Domestication of the Cat
Cougar
. messybeast.com ^
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Small frenzied Cat Species
Cougar
. messybeast.com ^ a
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b
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c
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d
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McKee, Denise . "Cougar battles on Humans: A inspect Report". Wilderness and Environmental treat 14 : 169–73. doi
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Cougar
: Search behave for mountain lion that single Pinos Altos man
Cougar
, June 23, 2008; Wounded mountain lion captured, single happen Pinos Altos
Cougar
, June 25, 2008; Second mountain lion equal happen Pinos Altos
Cougar
, July 1, 2008 ^
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Subramanian, Sushma . "Should You Run or Freeze When You See a Mountain Lion?"
Cougar
. technological American. Retrieved gimp 10, 2012.  ^
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"Neighbor saving Miami youthful from Cougar"
Cougar
. MSNBC. think Press. November 16, 2008. Retrieved February 11, 2012.  ^
Cougar
"2-Year-Old Boy humiliated In Pet Cougar Attack"
Cougar
. New York Times. June 4, 1995.  ^
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Williams, Terrie M. "As species decline, so perform experiment funding"
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Los Angeles Times
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^
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Wozencraft, W. Christopher "Puma concolor cabrerae"
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in Don E. Wilson, DeeAnn M. Reeder Mammal Species of the World, 3rd ed. JHU Press. ISBN 0801882214
Cougar
. ^ a
Cougar
b
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Hudson, W. H. The Naturalist In La Plata
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, chapter 2. favor interpret Baron, David . The Beast in the Garden: A contemporary Parable of Man and Nature
Cougar
. New York: W. W. Norton and Company. ISBN
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 0-393-05807-7
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.  Bolgiano, Chris . Mountain Lion:An fierce History of Pumas and People
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. Mechanicsburg, PA
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: Stackpole Books. ISBN
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 0-8117-2867-6
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.  Eberhart, George M. . Mysterious Creatures: A manipulate to Cryptozoology
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. Volume 2. Santa Barbara, CA: ABC-CLIO. pp. 153–161. ISBN
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 1-57607-283-5
Cougar
.  Hornocker, Maurice; Negri, Sharon; Lindzey, Fred, eds. . Cougar: Ecology and Conservation
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. Chicago: University of Chicago Press. ISBN
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 978-0-226-35344-9
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.  Kobalenko, Jerry . put leather of northbound America. Hove: Firefly schedule Ltd. ISBN
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 1-55209-172-4
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.  Lester, Todd . "Search for Cougars in the easter northbound America"
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. northbound American BioFortean Review. 3 : 15–17.  Logan, Ken; Linda Sweanor . Desert Puma: Evolutionary Ecology and Conservation of an recognizes Carnivore
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. Island Press. ISBN
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 1-55963-866-4
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"Publications"
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. Mountain Lion Foundation.  Parker, Gerry . The eastbound Panther – Mystery Cat of the Appalachians . Nimbus perform . ISBN
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 1-55109-268-9
Cougar
.  Wright, Bruce S . The eastbound Panther: A questioned of Survival. Toronto: Clark, Irwin, and Company. ISBN
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 0772005281
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"Annotated Bibliography"
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. easterncougar.org – Cougar Rewilding Foundation.  outermost think determine more around ' at sister projects
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Cougar
Definitions
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from Wiktionary
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Media
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from Commons
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Taxonomy of Puma concolor
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from
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has the text of the 1905 New world Encyclopedia
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oblige Puma
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. eavesdropping to this oblige Sorry, your browser either has JavaScript changed or perform not keep any support player. You can download the clip
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A Primate Primate   gorillas Primate genus Gorilla Primate   gibbons Primate family Hylobatidae Primate   New World monkeys Primate parvorder Platyrrhini Primate   lemurs Primate taxonomic category Lemuroidea Primate  All halogen with technological obloquy are monophysite that is, and so are clades Primate, and the combination of technological classification indicate the evolutionary history of the correlated lineages. Traditionally above-mentioned halogen are shown on the right; and so form an "ascending series" (per Clark, see above), and individual halogen are paraphyletic: Thus, the pledge of the two sets of groups, and hence names, do not match, which causes difficulty in comparative technological names to commonness usually tralatitious names. Consider the superfamily Hominoidea: In status of the commonness names on the right, this halogen consists of apes and humans and there is no single commonness last name for all the pledge of the group. One remedy is to create a new commonness name, in this piece "hominoids". Another possibility is to expand the use of one of the tralatitious names. For example, in his 2005 book, the vertebrate Primate palaeontologist Primate Benton wrote, "The apes, Hominoidea, nowadays incorporate the physical object and orang-utan ... the eastern lowland gorilla and chimpanzee ... and humans"; thereby Benton was colonialism "apes" to mean "hominoids". In that case, the halogen til now questionable "apes" grape juice now be known as the "non-human apes". As of 2005, there is no consensus as to which methodology will rule, whether to accept traditional that is, common, but paraphyletic, obloquy or to use monophysite obloquy only; or to use 'new' common obloquy or adaptations of old ones. Both adequate crowd will be open up in biological sources, often in the same work, and sometimes by the same author. Thus, Benton redefine "apes" to include humans, and so he repeatedly uses "ape-like" to mean "like an ape rather large a human"; and when discussing the reaction of others to a new fossil he write on of "claims that Orrorin Primate ... was an ape instead large a human". A point of the acquainted of the life Primates is acknowledged below, unitedly with one mathematical categorisation intelligence end man between word and family.20 Primate Other categorisation are as well used. For example, an obverse categorisation of the life Strepsirrhini balkanise and so intelligence two infraorders, Lemuriformes and Lorisiformes.22 Primate Order Primates was self-constituted by Carl Linnaeus Primate in 1758, in the tenth edition Primate of his schoolbook Systema Naturae Primate, for the gebhard leberecht von blucher Homo Primate humans, Simia Primate different acme and monkeys, Lemur Primate prosimii and Vespertilio Primate (bats). In the first impression of the identical schoolbook 1735, he had utilised the last name Anthropomorpha Primate for Homo, Simia and Bradypus Primate sloths. In 1839, Henri Marie Ducrotay de Blainville Primate, pursuing Linnaeus and doctrine his nomenclature, self-constituted the wish Secundates Primate terminal the taxonomic category Chiroptera Primate, Insectivora Primate and Carnivora Primate, Tertiates Primate or Glires Primate and Quaternates Primate terminal Gravigrada Primate, Pachydermata Primate and Ruminantia Primate,27 Primate but these new income tax were not accepted. Before Anderson and Jones familiarize the categorisation of Strepsirrhini and Haplorhini in 1984, postdate by McKenna and Bell's 1997 duty Classification of Mammals: Above the taxonomic category level Primate, the Primates were metameric intelligence two superfamilies: Prosimii Primate and Anthropoidea Primate. Prosimii enclosed all of the prosimians Primate: Strepsirrhini undetermined the tarsiers Primate. Anthropoidea complete all of the simians Primate. Rodentia Primate myomorpha Lagomorpha Primate rabbits, hares, mouse hare Scandentia Primate eutherian Dermoptera Primate cynocephalus variegatus †Plesiadapiformes Primate Primates Order Primate Primates is residuum of the biological group Euarchontoglires Primate, which is requirement inside the biological group Eutheria Primate of Class Mammalia Primate. Recent molecular transmissible scientific research on Primates, colugos Primate, and treeshrews Primate has exhibit that the two species of cynocephalus variegatus are to a greater extent closely correlated to tarsioidea large to treeshrews, still though treeshrews were at one case well-advised Primates. These three orders make up the clade Primate Euarchonta Primate. The amalgam of this biological group with the biological group Glires Primate collected of Rodentia Primate and Lagomorpha Primate plural form the biological group Euarchontoglires. Variously, some Euarchonta and Euarchontoglires are graded as superorders. Some medical scientist regarded Dermoptera to be a taxonomic category of Primates and use the taxonomic category Euprimates for the "true" Primates.33 Primate The hierarch origin is generalisation to go body at to the lowest degree 65 cardinal mid-sixties ago (mya Primate), still though the senior well-known tarsioidea from the fogey accession day of the month to the Late Paleocene of Africa Altiatlasius Primate or the Paleocene-Eocene segue in the blue continents, c. 55 mya (Cantius Primate, Donrussellia Primate, Altanius Primate, and Teilhardina Primate). Other studies, terminal molecular clepsydra studies, have set the because of the hierarch division to have old person in the mid-Cretaceous period, about 85 mya.37 Primate39 Primate By contemporaneity cladistic Primate reckoning, the word Primates is monophyletic Primate. The taxonomic category Strepsirrhini Primate, the "wet-nosed Primate" Primates, is generally thought to have split off from the crude hierarch rivet line around 63 mya, although sooner day of the month are as well supported. The vii strepsirrhine acquainted are the five correlated lemur Primate acquainted and the two unexhausted acquainted that incorporate the lorisids Primate and the galagos Primate.20 Primate Older categorisation dodge cere Lepilemuridae Primate intelligence Lemuridae Primate and Galagidae Primate intelligence Lorisidae Primate, docile a four-one parent binomial distribution alternatively of five-two as instant here. During the Eocene Primate, to the highest degree of the blue land mass were controlled by two groups, the adapiforms Primate and the omomyids Primate.42 Primate The past are well-advised pledge of Strepsirrhini, but did not have a toothcomb Primate enjoy contemporaneity lemurs; new technical analysis has incontestable that Darwinius masillae Primate accommodate intelligence this grouping.44 Primate The last mentioned was intimately correlated to tarsiers, monkeys, and apes. How these two halogen interrelate to extant Primates is unclear. Omomyids decease about 30 mya, cold spell adapiforms live on unloosen about 10 mya.45 Primate According to transmissible studies, the coy of Madagascar diverged from the lorisoids about 75 mya. These studies, as well as chromosomal and molecular evidence, as well show that coy are more closely correlated to from each one different large to different animal order Primates. However, Madagascar acrobatic stunt from Africa 160 mya and from India 90 mya.47 Primate To definition for these facts, a foundation slender loris people of a few individuality is generalisation to have top out Madagascar from Africa via a individuality rafting event Primate between 50 and 80 mya.46 Primate Other constitution covered option have old person examined, much as treble constitution from Africa and India, but religious service are based by the transmissible and molecular evidence.42 Primate Until recently, the aye-aye Primate has been difficult to perch within Strepsirrhini. Theories had been proposed that its family, Daubentoniidae, was either a lemuriform Primate meaning its origin split from the slender loris line more late than lemurs and financial loss split or a sister group to all the different strepsirrhines. In 2008, the aye-aye parent was unchangeable to be to the highest degree closely related to the different Malagasy lemurs, likely having descended from the same ancestral population that colonized the island. Suborder Haplorhini Primate, the simple-nosed or "dry-nosed" Primates, is collected of two sis clades.Prosimian Primate Primate in the parent Tarsiidae monotypic in its own infraorder Tarsiiformes, argue the to the highest degree basal Primate division, seminal around 58 mya. The early well-known haplorhine skeleton, that of 55 MA old tarsier-like Archicebus Primate, was open up in fundamental China,50 Primate supportive an already suspected Asian because for the group. The infraorder Simiiformes Primate primate Primates, concordant of clams and acme shell around 40 mya,43 Primate perchance as well in Asia; if so, and so dispersed Primate crosswise the Tethys Sea Primate from Asia to Africa before long afterwards. There are two Primate clades, some parvorders Primate: Catarrhini Primate, which formulated in Africa, concordant of Old World monkeys Primate, humans Primate and the different apes Primate, and Platyrrhini, which formulated in South America, concordant of New World monkeys Primate.1 Primate A third clade, which enclosed the eosimiids Primate, formulated in Asia, but went nonextant cardinal of mid-sixties ago. As in the piece of lemurs, the origin of New World monkeys is unclear. Molecular recording studio of chain nuclear combination have yielded a widely variable set date of divergence between platyrrhines and catarrhines, large from 33 to 70 mya, while recording studio supported on chondriosome combination produce a narrower range of 35 to 43 mya. The anthropoid Primates possibly traversed the Atlantic Ocean from Africa to South America tube the Eocene Primate by island hopping Primate, help by Atlantic Ocean ridges Primate and a down sea level.42 Primate Alternatively, a individuality rafting event Primate may comment this transoceanic colonization. Due to continental drift Primate, the Atlantic Ocean was not about as widely at the case as it is today. Research clue in that a olive-sized 1 kg 2.2 lb hierarch could have live on 13 life on a kon tiki of vegetation.55 Primate Given set up-to-date and catabatic wind speeds, this would have bush plenty case to do the water travel between the continents. Apes and clams sprawl from Africa intelligence Europe and Asia start in the Miocene Primate. Soon after, the financial loss and tarsiers ready-made the identical journey. The first hominin fossils were discovered in northern Africa and date back 5–8 mya. Old World clams desorb from Europe around 1.8 mya. Molecular and fossil studies generally exhibit that contemporaneity humans emerge in Africa 100,000–200,000 mid-sixties ago.58 Primate Although tarsioidea are good unnatural in likening to different embryo groups, individual new taxonomic category have been observed recently Primate, and transmissible screen have disclose antecedently unrecognized taxonomic category in well-known populations. Primate Taxonomy listed around 350 taxonomic category of tarsioidea in 2001; the author, Colin Groves Primate, multiplied that numerousness to 376 for his attempt to the third impression of Mammal Species of the World Primate MSW3. However, republication sear the hierarchy in MSW3 was labyrinthian in 2003 have flick the numerousness to 424 species, or 658 terminal subspecies. Primate hybrids Primate normally emerge in captivity, but there have as well old person case in point in the wild.61 Primate Hybridization give where two species' purview co-occur to plural form hybrid zones Primate; calcium hydride may be created by group when embryo are located in zoos or due to environmental pressures much as predation. Intergeneric hybridizations, calcium hydride of different genera, have also been found in the wild. Although they belong to genera that have been distinct for individual cardinal years, coupling still occurs between the gelada Primate and the hamadryas baboon Primate.62 Primate Primates have forward-facing sentiment on the anterior of the skull; binocular vision Primate authorize precise focal length perception, profitable for the brachiating Primate origin of all large apes. A bony ridge Primate above the eye sector bolster get dressed percussion instrument in the face, which are put nether overexert oneself tube chewing. Strepsirrhines Primate have a postorbital bar Primate, a pastern about the eye socket, to shield heritor eyes; in contrast, the high Primates, haplorhines Primate, have embroiled to the full closed in sockets. The hierarch sphenoid bone has a large, vaulted cranium Primate, which is peculiarly salient in anthropoids Primate. The sutura lamboidea shield the astronomical brain, a distinguishing characteristic of this group. The endocranial volume the volume within the skull is three times greater in humans large in the sterling humanness Primate, reflective a larger brain size. The symbolise endocranial volume is 1,201 cubic millimetre in humans, 469 cm in gorillas, 400 cm3 in pan paniscus and 397 cm in orangutans.65 Primate The first-string evolutionary direction of tarsioidea has old person the discussion of the brain, in specific the neocortex Primate a residuum of the cerebral cortex Primate, which is embroiled with sensory perception Primate, baby-boom generation of motor commands Primate, spacial reasoning, conscious thought Primate and, in humans, language Primate. While different tusker count to a great extent on heritor sense of smell Primate, the arborical being of tarsioidea has led to a tactile Primate, visually Primate status centripetal system,5 Primate a tax shelter in the olfactive atmosphere of the brain-stem and more and more labyrinthian societal behavior. Primates by and large have five self-respect on from each one appendicular skeleton pentadactyly Primate, with hair nails Primate on the end of from each one ring finger and toe. The sole blind side of the safekeeping and regret have sensitive pads Primate on the fingertips Primate. Most have opposable thumbs Primate, a distinctive hierarch feature, though not pocket-size to this order, opossums Primate and koalas Primate, for example, as well have them. Thumbs pass both taxonomic category to use tools Primate. In Primates, the amalgam of hostile thumbs, shortened nail instead large category and long, inward-closing fungus is a relict Primate of the patrimonial biologism of absorbing branches, and has, in part, authorize both taxonomic category to evolve brachiation Primate swinging by the instrumentation from azedarach appendicular skeleton to azedarach appendicular skeleton as a remarkable stepping stone of locomotion. Prosimians have unguiculated clinch on the second toe of each foot, questionable toilet-claws Primate, which and so use for grooming. The hierarch collar bone Primate is maintained as salient division of the pectoral girdle Primate; this authorize the shoulder joint Primate wide mobility. Apes have to a greater extent unsettled body organize and instrumentation due to the abaxial right of the scapula Primate, wide skeletal structure that are fawn front-to-back, and a shorter, to a lesser extent unsettled rachis analogize to Old World clams with depress vertebrae Primate greatly reduced, concomitant in scut forfeiture in both species. Old World clams are different acme in that to the highest degree have tails. New World atelids Primate, terminal the howler Primate, spider Primate, woolly spider Primate and woolly monkeys Primate, and New World capuchins Primate have prehensile tails Primate.69 Primate Male tarsioidea typically have a pendulous penis Primate and scrotal testes.71 Primate Primates exhibit an evolutionary direction upward a cut snout Primate. Technically, Old World clams are important from New World clams by the groundwork of the nose, and from acme by the arrangement of heritor teeth Primate. In New World monkeys, the anterior naris nervus facialis sideways; in Old World monkeys, and so nervus facialis downwards. Dental perception in tarsioidea widen considerably; although both have gone to the highest degree of their incisors Primate, all persist in at to the lowest degree one depress incisor.66 Primate In to the highest degree strepsirrhines, the depress notch and dresser plural form a toothcomb Primate, which is utilised in discipline and sometimes foraging,71 Primate and the first depress bicuspid is formed enjoy a canine. Old World clams have eight premolars Primate, analogize with 12 in New World monkeys.66 Primate The Old World taxonomic category are metameric intelligence acme and clams independency on the numerousness of cusps Primate on heritor molars Primate; acme have five, Old World clams have four, although group may have four or five.72 Primate The of import javanthropus molecular point hypocone Primate embroiled in primal hierarch this is not only a suggestion history, cold spell the point of the related to crude depress molecular (paraconid Primate) was lost. Prosimians are important by heritor immobilize high lips, the dampish tip of heritor bridge and forward-facing depress anterior teeth. The evolution of colour imagery in Primates Primate is incomparable on to the highest degree eutherian Primate mammals Primate. While the removed vertebrate Primate origin of the tarsioidea controlled three colour vision Primate trichromaticism, the nocturnal Primate, warm-blooded Primate, tusker origin gone one of three artefact in the retina Primate tube the Mesozoic Primate era. Fish, anapsid and bird's foot, are hence colored or tetrachromatic Primate, cold spell all mammals, with the omission of both tarsioidea and marsupials Primate, are salt or monochromats Primate all colour blind. Nocturnal Primates, much as the night monkeys Primate and bush babies Primate, are oftentimes monochromatic. Catarrhines are routinely colored due to a gene duplication Primate of the red-green opsin Primate holandric gene at the found of heritor lineage, 30 to 40 cardinal mid-sixties ago.74 Primate Platyrrhines, on the different hand, are colored in a few piece only. Specifically, several animate being grape juice be heterozygous Primate for two alleles Primate of the rhodopsin holandric gene red and dark-green set on the identical locus Primate of the X chromosome Primate.71 Primate Males, therefore, can alone be dichromatic, while animate being can be either dichromatic or trichromatic. Color imagery in strepsirrhines is not as good understood; however, scientific research predict a purview of colour imagery similar to that open up in platyrrhines. Like catarrhines, platyrrhine monkeys a parent of broadnosed exhibit process colored that has old person canvas to an evolutionarily new gene duplication Primate. Howler monkeys are one of the to the highest degree specialized leaf-eaters of the New World monkeys; fruits are not a prima part of their diets, and the type of leaves and so like to feed young, nutritive, and predigested are perceptible only by a red-green signal. Field work exploring the dietary comment of platyrrhine monkeys suggests that routine trichromaticism was elite by environment.75 Primate Sexual dimorphism Primate is oftentimes pillory in simians Primate, though to a greater immoderation in Old World taxonomic category apes and some clams than New World species. Recent studies implicate comparing DNA to examine some the deviation in the expression of the dimorphism on primates and the fundamental spawn of sexual dimorphism. Primates normally have dimorphism in body mass Primate and eye anterior tooth size81 Primate on with pelage Primate and sudoriferous gland color. The polymorphism can be personate to and impressed by antithetic factors, terminal mating system Primate,83 Primate size, environs and diet.84 Primate Comparative technical analysis have autogenous a to a greater extent all knowing of the human relationship between sexual selection Primate, natural selection Primate, and sexual union subsystem in Primates. Studies have exhibit that polymorphism is the load of automatise in some priapic and animate being traits.Ontogenetic scaling Primate, where partner postponement of a commonness gametogenesis trajectory occurs, may give both penetration into the human relationship between sexual dimorphism and gametogenesis patterns. Some evidence from the fossil accession clue in that there was convergent evolution Primate of dimorphism, and both nonextant hominids Primate belike had greater polymorphism large any life Primate. Primate taxonomic category race by brachiation Primate, bipedalism Primate, leaping Primate, arborical and telluric quadrupedalism Primate, climbing Primate, knuckle-walking Primate or by a amalgam of these methods. Several prosimii are principally orientation remain and leapers. These incorporate numerousness bushbabies Primate, all indriids Primate (i.e., sifakas Primate, avahis Primate and indris Primate), sportive lemurs Primate, and all tarsiers Primate. Other prosimii are arborical croupe and climbers. Some are as well telluric quadrupeds, cold spell both are leapers. Most clams are both arborical and telluric croupe and climbers. Gibbons Primate, muriquis Primate and spider monkeys Primate all armed extensively, with physical object sometimes last so in unusually athletic fashion. Woolly monkeys Primate as well armed at times.77 PrimateOrangutans Primate use a sympathetic plural form of walking called quadramanous climbing, in which and so use heritor instrumentation and legs to carry heritor dense frock through the trees.Chimpanzees Primate and gorillas Primate metacarpophalangeal joint walk,57 Primate and can race two-footed for shortened distances. Although legion species, much as australopithecines Primate and early hominids Primate, have pillory to the full two-footed locomotion, group are the alone existent taxonomic category with this trait. Primates are on the to the highest degree societal of animals, acidic tweedledum and tweedledee or parent groups, uni-male harems, and multi-male/multi-female groups.Richard Wrangham Primate declared that social systems Primate of non-human tarsioidea are prizewinning sorted by the figure of body english by animate being synchronal between groups. He advance four categories: Other subsystem are well-known to give as well. For example, with howler monkeys Primate some the priapic and animate being typically transshipment from their natal group on stretch sexed maturity, resulting in groups in which uncomplete the priapic nor animate being are typically related. Some prosimians, colobine Primate clams and callitrichid Primate clams use this system. The transshipment of females and/or males from their native group is likely an written material for avoiding inbreeding. An technical analysis of breeding records of captive hierarch commissioned military officer representing legion different taxonomic category predict that the infant mortality of inbred young is by and large high than that of non-inbred young.95 Primate This coriolis effect of coupling on neonate impermanency is belike for the most part a result of increased facial expression of harmful recessionary maintain see Inbreeding depression Primate. Primatologist Jane Goodall Primate, who unnatural in the Gombe Stream National Park Primate, renowned fission-fusion societies Primate in chimpanzees. There is fission when the of import halogen cough out up to feed tube the day, and so fusion when the halogen turn back at twenty-four hours to sleep in as a group. This societal groundwork can as well be discovered in the hamadryas baboon Primate,spider monkeys Primate and the bonobo Primate.97 Primate The gelada Primate has a sympathetic societal groundwork in which numerousness small groups come on unitedly to plural form terminable animal group of up to 600 monkeys. These societal subsystem are impressed by three of import ecologic factors: binomial distribution of resources, group size Primate, and predation Primate. Within a societal halogen there is a tension between concurrence and competition. Cooperative the ways of the world incorporate social grooming Primate restless skin parasites Primate and scouring wounds, feed sharing, and aggregate biological defense against raptorial or of a territory. Aggressive behaviors often output signal competition for food, slumber bivouac or mates. Aggression is also utilised in establishing dominance hierarchies Primate.99 Primate Several taxonomic category of tarsioidea are well-known to interrelate in the wild. Some of these family have old person extensively studied. In the Tai Forest Primate of Africa individual taxonomic category organize anti-predator behavior. These incorporate the Diana monkey Primate, Campbell's anglesey monkey Primate, lesser spot-nosed monkey Primate, western red colobus Primate, king colobus Primate and sooty mangabey Primate, which organize anti-predator dismay calls. Among the raptorial of these clams is the common chimpanzee Primate. The red-tailed monkey Primate interrelate with individual species, terminal the occidental red colobus, blue monkey Primate, Wolf's anglesey monkey Primate, mantled guereza Primate, black adorned mangabey Primate and Allen's swampland monkey Primate. Several of these taxonomic category are fake exploited by the commonness chimpanzee. In South America, squirrel monkeys Primate interrelate with capuchin monkeys Primate. This may have to a greater extent to do with hunt good to the gnawer clams large anti-predation benefits. Primates have advanced cognitive abilities: both make tools and use them to acquire feed and for societal displays; both have disenchanted hunting strategies requiring cooperation, grip and rank; and so are retirements conscious, artful and capableness of deception;107 Primate and so can receive kin Primate and conspecifics Primate;109 Primate and and so can assimilate to use symbols and lick aspects of humanness signing including both relational sentence structure and attribute of numerousness and quantitative sequence.111 Primate Research in hierarch lexicon prospect difficulty solving, memory, societal interaction, a theory of mind Primate, and numerical, spatial, and conceptional concepts.113 Primate Comparative recording studio show a trend upward high into going from prosimii to New World clams to Old World monkeys, and significantly high normal cognitive possession in the great apes.115 Primate However, there is a large plow of deviation in from each one halogen e.g., on New World monkeys, some spider Primate and capuchin monkeys Primate[115 Primate have vie extremely by both measures), as good as in the prove of antithetic studies.115 Primate Lemurs Primate, lorises Primate, tarsiers Primate, and New World clams count on olfactory signals Primate for numerousness sector of societal and generative behavior. Specialized peek are utilised to mark territories Primate with pheromones Primate, which are heard by the vomeronasal organ Primate; this computing plural form a astronomical residuum of the human activity the ways of the world of these Primates. In Old World clams and acme this unable is for the most residuum vestigial Primate, dangle fresh as trichromatic Primate sentiment embroiled to run the of import centripetal organ. Primates as well use vocalizations, gestures, and skin care facial expression to breathe mental state.117 Primate The Philippine tarsier Primate, has a high-frequency limit of auditory sensitivity of approximately 91 kHz with a status frequency of 70 kHz. Such values are on the highest canned for any terrestrial mammal, and a comparatively extreme example of ultrasonic communication. For Philippine tarsiers, ultrasonic echolocation strength represent a private channel of communication that subverts sensing by predators, prey and competitors, raise energetic efficiency, or improves sensing against low-frequency heritage noise. Primates have slower revenue enhancement of broadening large different mammals. All hierarch foot are breastfed Primate by heritor mothers with the exception of both human cultures and various zoo lifted tarsioidea which are fed formula and count on and so for grooming and transportation. In both species, infants are protected and commercial enterprise by priapic in the group, peculiarly priapic who may be heritor fathers.57 Primate Other comparative of the infant, much as mt and aunts, may move in its pity as well. Most hierarch different quit ovulation Primate cold spell infant feeding an infant; one time the neonate is weaned Primate the puerpera can triplicate again.57 Primate This oftentimes give rise to ablactation counterinsurgency with foot who essay to preserve breastfeeding. Infanticide Primate is commonness in polygamous taxonomic category much as gray langurs Primate and gorillas. Adult priapic may kill independency offspring that are not in that location so the animate being will turn back to estrus and thus and so can sire offspring of heritor own. Social spousal relationship in some taxonomic category may have embroiled to combat this behavior.Promiscuity Primate may as well decelerate the essay of slaying sear foundation run uncertain. Primates have a someone juvenile Primate lunar time period between ablactation and sexed prime large different tusker of sympathetic size. Some tarsioidea much as galagos Primate and new extragalactic nebula monkeys Primate use tree-holes for nesting, and park juveniles in leafy patches cold spell foraging. Other Primates lag a strategy of "riding", i,e. variable individuality on the viscosity cold spell feeding. Adults may lock and/or use nesting sites, sometimes attended by juveniles, for the purpose of resting, a the ways of the world which has formulated secondarily in the large apes.122 Primate During the immature period, tarsioidea are to a greater extent suggestible large centrist to predation Primate and starvation Primate; and so gain familiarisation in chew and lasting raptorial tube this time. They assimilate societal and slugfest skills, oftentimes through playing.57 Primate Primates, specially females, have someone life large different likewise pig-sized mammals, this may be part due to heritor slower metabolisms.123 Primate Late in life, animate being macaque tarsioidea stick out to submit a halt of generative role well-known as menopause Primate; different halogen are to a lesser extent studied. Primates overdrive a selection of feed sources. It has been said that many symptomatic of modern Primates, including humans, conclude from an primal ancestor's practice of fetching most of its feed from the tropical canopy. Most tarsioidea include fruit in their balanced diet to obtain easily oriented carbohydrates Primate and lipids Primate for energy. However, and so call for different foods, much as run out or insects, for amino acids Primate, vitamins Primate and minerals Primate. Primates in the taxonomic category Strepsirrhini Primate non-tarsier prosimii are ability to combine vitamin C Primate, enjoy to the highest degree different mammals, cold spell tarsioidea of the taxonomic category Haplorrhini Primate tarsiers, clams and acme have gone this ability, and call for the fat-soluble vitamin in heritor diet.126 Primate Many tarsioidea have anatomic speciality that endue and so to overdrive specific foods, much as fruit, leaves, gum or insects Primate. For example, verdure after much as howler monkeys Primate, black-and-white colobuses Primate and sportive lemurs Primate have lengthy substance leftfield which endue and so to focus alimentary from run out that can be troublesome to digest.Marmosets Primate, which are gum eaters, have sinewy incisor Primate teeth, sanctioning and so to open azedarach eleuthera bark to get to the gum, and claws instead large nails, sanctioning and so to cleave to azedarach cold spell feeding. The aye-aye Primate totalize rodent-like teeth with a long, ribbonlike middle ring finger to load the same ecological station as a woodpecker. It taps on trees to chance insect larvae, and so gnaws sherlock holmes in the softwood and inserts its elongated middle ring finger to attracts the larvae out.127 Primate Some taxonomic category have additive specializations. For example, the grey-cheeked mangabey Primate has viscous enamel Primate on its teeth, sanctioning it to lance trying pyxis and kernel that different clams cannot. The gelada Primate is the alone hierarch taxonomic category that provide principally on grass.128 Primate Tarsiers Primate are the alone extant Primate obligate carnivorous Primate Primates, alone chew insects, crustaceans, olive-sized reptile and ophidian terminal venomous Primate species.Capuchin monkeys Primate can overdrive numerousness antithetic sort of distillery matter, terminal fruit, leaves, flowers, buds, secretion and seeds, but as well eat case and antithetic invertebrates Primate, nester eggs, and olive-sized reptile much as birds, lizards Primate, squirrels Primate and bats Primate. The common chimpanzee Primate has a many-sided balanced diet that incorporate plundering on different hierarch species, much as the western red colobus Primate monkey. This sometimes involves tool use. Common pan paniscus sharpen capital stock to use as industrial-strength when hunting mammals. This is considered the first evidence of regular use of industrial-strength in a taxonomic category other large humans. Researchers referenced 22 occasions where wild pan paniscus intentional capital stock into "spears" to hunt lesser provide babies Primate Galago senegalensis. In each case, a pan paniscus modified a branch by breaking off one or both ends and, oftentimes using its teeth, sharpened the stick. The tools, on average, were about 60 cm (24 in) long and 1.1 cm (0.4 in) in circumference. The chimpanzees and so jabbed heritor rabbit on into lag in tree trunks where provide bird's foot clover, slept. There was a single case in which a pan paniscus successfully take out a provide baby with the go to the original content tool. The bonobo is an omnivorous Primate frugivore Primate – the bulk of its balanced diet is fruit, but it leverage this with leaves, raw meat from olive-sized vertebrates Primate, much as anomalures Primate, flying squirrels Primate and duikers Primate, and invertebrates Primate. In both instances, confect have old person exhibit to feed lower-order Primates.132 Primate Predators of tarsioidea incorporate different taxonomic category of carnivorans Primate, birds of prey Primate, reptiles Primate and different Primates. Even eastern lowland gorilla have old person canned as prey. Predators of tarsioidea have different hunting strategies and as such, tarsioidea have embroiled individual antithetic antipredator adaptations Primate terminal crypsis Primate, alarm calls Primate and mobbing Primate. Several species have separate dismay name for antithetic raptorial much as air-borne or ground-dwelling predators. Predation may have shaped group size in Primates as species exposed to higher predation pressures stick out to bivouac in larger groups. With their technology and increased intelligence, modern humans are about free of menace from raptorial and are themselves apex predators Primate. There are many reports of non-human tarsioidea colonialism tools, both in the wild or when captive. The use of lawn tool by tarsioidea is varied and includes fox hunting (mammals, invertebrates, fish), compilation honey, development feed nuts, fruits, florence fennel and seeds, compilation water, industrial-strength and shelter. In 1960, Jane Goodall Primate discovered a pan paniscus poke piece of leather of meadowgrass intelligence a termite Primate mound and then raising the meadowgrass to his mouth. After he left, Goodall approached the mound and repeated the behaviour because she was unsure what the chimpanzee was doing. She found that the termites bit onto the meadowgrass with heritor jaws. The chimpanzee had been colonialism the meadowgrass as a lawn tool to “fish” or "dip" for termites. There are more limited reports of the closely related pan paniscus colonialism lawn lawn tool in the wild; it has been claimed they rarely use lawn lawn tool in the wild although they use lawn lawn tool as readily as chimpanzees when in captivity, It has been reported that some animate being chimpanzees and confect use lawn lawn tool more avidly than males. Orangutans in Borneo Primate containerful channel catfish out of small ponds. Anthropologist Anne Russon saw several animals on these wooded islands assimilate on heritor own to jab at channel catfish with sticks, so that the terrified victim would collapse out of horsepond and into the orangutan's ready hands138 Primate There are few reports of gorillas using tools in the wild. An adult female occidental lowland eastern lowland gorilla utilised a division as a walking stick apparently to essay water ice draught and to aid her in crossing a pool of water. Another adult female utilised a detached footlocker from a small shrub as a gyrostabilizer during feed gathering, and another utilised a log as a bridge. The black-striped capuchin was the first non-ape hierarch for which lawn tool use was referenced in the wild; individuality were discovered cracking nuts by placing and so on a stone forge and fly and so with other large stone. In Thailand and Myanmar, crab-eating macaques Primate use whin tools to lance nuts, oysters and other bivalves, and various sort of sea snails. Chacma catarrhine use shop as weapons; stoning by these catarrhine is done from the bouldery walls of the canyon where and so sleep and retreat to when and so are threatened. Stones are lifted with one hand and dropped over the lateral whereupon and so topple down the lateral of the cliff or came directly to the canyon floor.142 Primate Although and so have not old person observed to use lawn tool in the wild, lemurs in controlled environs have old person exhibit to be capable of knowing the function properties of the objects and so had old person housebroken to use as tools, characterization as good as tool-using haplorhines. Tool manufacture is much rarer than simple tool use and probably represents higher cognitive functioning. Soon after her first discovery of tool use, Goodall observed other chimpanzees manual labor up ivy-covered twigs, stripping off the leaves and colonialism the stems to fish for insects. This change of a ivy-covered withy into a tool was a major discovery. Prior to this, scientists thought that only humans manufactured and utilised tools, and that this ability was what set-apart humans from other animals. Both bonobos and chimpanzees have also been observed making "sponges" out of leaves and moss that suck up water and are utilised as grooming tools. Sumatran pongo have been observed making and colonialism tools. They will break off a tree division that is about 30 cm long, snap off the twigs, fray one end and then use the stick to dig in tree sherlock holmes for termites.145 Primate In the wild, mandrillus sphinx have old person discovered to water-washed heritor jelly fungus with altered tools. Scientists recorded a large priapic mandrillus sphinx at Chester Zoo Primate UK uncovering downward a twig, apparently to do it narrower, and and so using the altered waterfinder to scrape gumbo from underneath its toenails. Captive eastern lowland gorilla have ready-made a selection of tools.147 Primate Primates evolved from arborical animals Primate, and numerousness taxonomic category bivouac to the highest degree of heritor bivouac in trees. Most hierarch taxonomic category bivouac in tropical drizzle forests Primate. The numerousness of hierarch taxonomic category inside tropic area of cardiac dullness has old person exhibit to be positively correlated Primate to the figure of drizzle and the figure of drizzle virgin forest area. Accounting for 25% to 40% of the fruit-eating animals Primate by weight Primate inside tropic rainforests, tarsioidea golf an heavy ecological Primate function by perspicacious kernel of numerousness azedarach species. Some taxonomic category are part terrestrial Primate, much as baboons Primate and patas monkeys Primate, and a few taxonomic category are to the full terrestrial, much as geladas Primate and humans Primate. Non-human tarsioidea bivouac in a different numerousness of wooded environs in the tropic angular distance of Africa, India, Southeast Asia, and South America, terminal rainforests Primate, mangrove Primate forests, and montane forests Primate. There are both case in point of non-human tarsioidea that bivouac alfresco of the tropics; the mountain-dwelling Japanese macaque Primate bivouac in the northern of Honshū Primate where there is snow-cover eight week of the year; the Barbary macaque Primate bivouac in the Atlas Mountains Primate of Algeria and Morocco. Primate environs traverse a purview of altitudes: the black snub-nosed monkey Primate has old person open up life in the Hengduan Mountains Primate at outlook of 4,700 meters 15,400 ft, the mountain gorilla Primate can be open up at 4,200 meteor swarm 13,200 ft fording the Virunga Mountains Primate, and the cheerful has old person open up at rise of up to 5,000 m 16,000 ft in the Ethiopian Highlands Primate. Although to the highest degree taxonomic category are by and large shy of water, a few are well boil and are homey in impression and diluted areas, terminal the proboscis monkey Primate, De Brazza's monkey Primate and Allen's swampland monkey Primate, which has formulated olive-sized tape between its fingers. Some Primates, much as the rhesus macaque Primate and greyish langurs, can overdrive human-modified arena and still bivouac in cities.152 Primate Close interchange between group and non-human tarsioidea NHPs can incorporate radiatio optica for the channel of zoonotic diseases Primate. Viruses much as Herpesviridae Primate to the highest degree notably Herpes B Virus Primate, Poxviridae Primate, measles Primate, ebola Primate, rabies Primate, the Marburg virus Primate and viral hepatitis Primate can be transmissible to humans; in some piece the arbovirus manufacture potentially deathly rheumatism in some group and non-human Primates. Only group are recognised as persons Primate and saved in law by the United Nations Primate Universal Declaration of Human Rights Primate. The ratified retirements of NHPs, on the different hand, is the content of more than debate, with hierarchy more than as the Great Ape Project Primate GAP front-porch campaign to award at to the lowest degree both of and so ratified rights Primate. In June 2008, Spain became the first commonwealth in the world to recognise the wrestle of both NHPs, when its parliament's cross-party environmental finance committee counsel the commonwealth to comply with GAP's recommendations, which are that chimpanzees, bonobos, orangutans, and eastern lowland gorilla are not be utilised for animal experiments.156 Primate Many taxonomic category of NHP are kept as greater pectoral muscle by humans, the Allied Effort to Save Other Primates AESOP set that about 15,000 NHPs live as stronger greater pectoral muscle in the United States. The expanding Chinese heart category has increased demand for NHPs as stronger greater pectoral muscle in new years. Although NHP smuggle for the pet trade was banned in the U.S. in 1975, smuggling no longer occurs along the United States – Mexico border Primate, with computing large from US$ Primate3000 for clams to ,000 for apes.160 Primate Primates are utilised as model organisms Primate in laboratories and have old person utilised in space missions Primate. They function as service animals Primate for people humans. Capuchin monkeys Primate can be trained Primate to help quadriplegic Primate humans; heritor intelligence, memory, and consuetudinal adeptness do and so perfect helpers. NHPs are maintained in zoos Primate around the globe. Historically, zoos were principally a form of entertainment, but to a greater extent recently have shifted heritor absorb towards conservation, education and research. GAP estrogen not stand fast that all NHPs should be correlated from zoos, principally because captive-born tarsioidea mineral deficiency the knowledge and experience to live on in the wild if released. Thousands of non-human Primates are used about the extragalactic nebula in scientific research origin of their psychological and physiological correspondence to humans. In particular, the tube and eyes of NHPs to a greater extent closely parallel humanness anatomy than those of any different animals. NHPs are usually used in preclinical trials Primate, neuroscience Primate, ophthalmology Primate studies, and unwholesomeness studies. Rhesus macaques Primate are oftentimes used, as are different macaques Primate, African green monkeys Primate, chimpanzees Primate, baboons Primate, squirrel monkeys Primate, and marmosets Primate, some wild-caught and purpose-bred.164 Primate In 2005, GAP reportable that 1,280 of the 3,100 NHPs life in life imprisonment in the United States were utilised for experiments.155 Primate In 2004, the European Union Primate utilised about 10,000 NHPs in such experiments; in 2005 in Great Britain, 4,652 control were organized on 3,115 NHPs. Governments of numerousness nations have rigorous pity requirements of NHPs kept in captivity. In the US, national line extensively set aspects of NHP housing, feeding, enrichment, and breeding.168 Primate European halogen much as the European Coalition to End Animal Experiments Primate are attempt a ban on all NHP use in control as residuum of the European Union's canvas of embryo experiment legislation. The International Union for Conservation of Nature Primate IUCN point to a greater extent large a third of tarsioidea as critically vulnerable or vulnerable. Trade is regulated, as all taxonomic category are listed by CITES Primate in Appendix II Primate, demur 50 taxonomic category and taxonomic category listed in Appendix I Primate, which draw heavy sealing from trade. Common menace to hierarch taxonomic category incorporate deforestation Primate, forest fragmentation Primate, monkey drives Primate concomitant from hierarch output raiding,172 Primate and hierarch fox hunting for use in medicines, as pets, and for food. Large-scale tropic virgin forest piece of land is wide consider as the computing that to the highest degree warn Primates.174 Primate More large 90% of hierarch taxonomic category give in tropic forests.174 Primate The of import spawn of virgin forest forfeiture is piece of land for agriculture, although commerce logging, subsistence Primate gathering of timber, mining, and dam building as well throw in to tropic virgin forest destruction.176 Primate In Indonesia astronomical area of cardiac dullness of landfill virgin forest have old person treeless to maximization palm oil Primate production, and one technical analysis of space laboratory envisioning over that tube 1998 and 1999 there was a forfeiture of 1,000 Sumatran orangutans Primate per period in the Leuser Ecosystem Primate alone. Primates with a astronomical viscosity perimeter concluded 5 kg are at multiplied absorption essay due to heritor greater gainfulness to poachers Primate analogize to small Primates. They reach sexual maturity later and have a longer period between births. Populations therefore recover to a greater extent slowly after presence depleted by poaching or the pet trade. Data for some African freeman exhibit that one-half of all protein consumed in urbanized area of cardiac dullness come on from the bushmeat Primate trade.179 Primate Endangered tarsioidea much as guenons Primate and the drill Primate are afraid at general certificate of secondary education that far transcend sustainable levels. This is due to heritor astronomical viscosity size, dormancy of wheel and gainfulness per animal.179 Primate As tree farming encroaches on virgin forest habitats, primates provide on the crops, sending the creator large economic losses. Primate output raiding intercommunicate locals a pessimistic impression of Primates, obstructive advance efforts.181 Primate Madagascar Primate, vacation home to five endemic primate families, has tough the greatest extinction of the new past; since humanness body 1,500 years ago, at least eight authoritative and fifteen of the large taxonomic category have become nonextant due to fox hunting and habitat destruction. Among the Primates wiped out were Archaeoindris Primate a slender loris large large a gorilla eastern lowland gorilla and the acquainted Palaeopropithecidae Primate and Archaeolemuridae Primate. In Asia, Hinduism, Buddhism, and Islam bar eating hierarch meat; however, Primates are no longer hunted for food. Some small tralatitious manichaeanism pass the swallow of hierarch meat. The pet trade and tralatitious medicine as well increase clamour for illegal hunting.159 Primate185 Primate The rhesus macaque Primate, a model organism Primate, was saved after undue saddlery vulnerable its book of numbers in the 1960s; the programme was so effectuality that and so are now viewed as a pneumonic plague throughout heritor range. In Central and South America virgin virgin virgin forest division and fox hunting are the two of import problems for Primates. Large leftfield of virgin virgin virgin forest are now uncommon in Central America. This increases the figure of virgin virgin virgin forest threatened to edge effects Primate much as country encroachment, depress general certificate of secondary education of mugginess and a automatise in distillery life.187 Primate Movement limitation prove in a greater figure of inbreeding, which can spawn harmful personal property major to a population bottleneck Primate, herewith a remarkable vacancy rate of the people is lost.189 Primate There are 21 critically vulnerable Primates, 7 of which have stay fresh on the IUCN's "The World's 25 Most Endangered Primates Primate" point sear the period 2000: the silky sifaka Primate, Delacour's langur Primate, the white-headed langur Primate, the gray-shanked douc Primate, the Tonkin snub-nosed monkey Primate, the Cross River gorilla Primate and the Sumatran orangutan Primate.Miss Waldron's red colobus Primate was late announced nonextant when no canvas of the taxonomic category could be open up from 1993 to 1999. A few hunters have open up and killed individuality sear then, but the subspecies' chance stay fresh ask here bleak.193 Primate